Page 2 of 7

BI9.1-3 | Minerals, electrolytes, Water and Acid base balance — Part 1

CLINICAL SCENARIO

Outpatient morning at a district hospital in Rajasthan. Three patients in consecutive beds:

Bed 1: A 22-year-old woman who is 28 weeks pregnant with pallor, fatigue, and dyspnoea on climbing stairs. Haemoglobin: 6.8 g/dL. Blood film: microcytic hypochromic red cells.

Bed 2: A 65-year-old man admitted after a fracture of the hip from a simple fall at home. DEXA scan shows T-score −3.2. He has been on long-term prednisolone for COPD.

Bed 3: A 45-year-old with severe diarrhoea and vomiting for 3 days. Blood gas: pH 7.28, pCO₂ 28 mmHg, HCO₃⁻ 13 mEq/L.

Three different mineral/electrolyte problems. Can you identify each one and explain the biochemistry?

WHY THIS MATTERS

Minerals, electrolytes, and acid-base balance are central to clinical practice at every level:

  • Iron deficiency anaemia affects 50% of pregnant women in India and is the leading cause of maternal mortality
  • Electrolyte disturbances (hypo/hypernatraemia, hypo/hyperkalaemia) are common in hospitalised patients and can be life-threatening
  • Acid-base disorders occur in every ICU patient and in common conditions: diabetic ketoacidosis, diarrhoea, renal failure, respiratory failure
  • Understanding calcium-phosphate metabolism underpins management of osteoporosis, CKD-mineral bone disease, and parathyroid disorders

These topics bridge Biochemistry directly to Medicine, Surgery, and Obstetrics — you will use them daily as a clinician.

RECALL

Recall from previous topics:

  • Haemoglobin structure: haem (iron-porphyrin) + globin chains — from Biochemistry (proteins)
  • Ca²⁺ and PTH/Vitamin D axis — from the Vitamins and Minerals chapter
  • Buffer concept from Chemistry: weak acid + its conjugate base
  • Kidney anatomy and filtration (from Anatomy/Physiology) — relevant to electrolyte regulation
  • Henderson-Hasselbalch equation (from general chemistry): pH = pKa + log ([A⁻]/[HA])

Iron — Absorption, Transport and Storage

Iron is the most clinically important mineral in India — deficiency is the leading nutritional disorder.

Iron — Absorption, Transport and Storage

Figure: Iron — Absorption, Transport and Storage

Multi-panel illustration of iron metabolism: duodenal absorption with DMT1 and ferroportin, hepcidin regulatory mechanism, body iron distribution and recycling, and haemochromatosis iron overload with organ damage

Total body iron: ~3.5–4 g (adult male); 2–3 g (adult female)
- ~65% in haemoglobin
- ~10% in myoglobin and enzymes (cytochromes, succinate dehydrogenase, ribonucleotide reductase)
- ~20–25% stored as ferritin and haemosiderin (liver, spleen, bone marrow)
- ~0.1% in plasma bound to transferrin (transport protein)

Iron absorption (duodenum and proximal jejunum):
- Haem iron (Fe²⁺ from meat, fish): absorbed directly via HCP1 receptor → much more bioavailable (~20–30%)
- Non-haem iron (Fe³⁺ from vegetables, cereals): must first be reduced to Fe²⁺ by brush border ferrireductase (Dcytb) — enhanced by Vitamin C; inhibited by phytates (cereals), oxalates (spinach), tannins (tea)
- Divalent metal transporter 1 (DMT1): transports Fe²⁺ into enterocyte
- Ferroportin: basolateral transporter → Fe²⁺ into portal blood → oxidised by hephaestin → Fe³⁺ → bound to transferrin (carries 2 Fe³⁺ per molecule)

Hepcidin — the master regulator of iron homeostasis:
- Peptide hormone made by the liver
- Binds ferroportin → internalises it → blocks Fe release from enterocytes, macrophages, hepatocytes
- High hepcidin → iron trapped → anaemia of chronic disease
- Low hepcidin → iron flows freely → used in haemochromatosis, iron overload

Iron storage:
- Ferritin: soluble storage protein; serum ferritin is the best marker of iron stores (low = iron deficiency; high = inflammation, iron overload, liver disease)
- Haemosiderin: insoluble ferritin aggregate in iron overload

Iron Deficiency Anaemia — Stages and Diagnosis

Iron Deficiency Anaemia vs Anaemia of Chronic Disease — Diagnostic Markers

Marker Iron Deficiency Anaemia (IDA) Anaemia of Chronic Disease (ACD)
Serum iron Low Low
TIBC (Total Iron Binding Capacity) High (body trying to capture more iron) Low or normal
Transferrin saturation Low (<20%) Low
Serum ferritin Low (<12 µg/L) — KEY distinguishing marker Normal or high (acute phase reactant)
Hepcidin Low (maximising absorption) High (IL-6 driven; traps iron in macrophages)
MCV Low (<80 fL) — microcytic Normal or low (normocytic or mildly microcytic)
Reticulocyte Hb content Low (early sensitive marker) Low or normal
Blood film Microcytic hypochromic; pencil cells, target cells Normocytic normochromic or mildly microcytic

Iron deficiency anaemia (IDA) develops in three stages:

Iron Deficiency Anaemia — Stages and Diagnosis

Figure: Iron Deficiency Anaemia — Stages and Diagnosis

Multi-panel illustration of iron deficiency anaemia: three progressive stages with biomarkers, diagnostic panel comparing IDA vs anaemia of chronic disease, blood film morphology showing microcytic hypochromic cells, and India prevalence with dietary causes

Stage 1 — Iron depletion: Iron stores (ferritin) depleted. No anaemia yet. Serum ferritin low (<12 µg/L).

Stage 2 — Iron-deficient erythropoiesis: Serum transferrin rises (body tries to capture more iron); transferrin saturation falls (<20%). Mild microcytosis. No significant anaemia.

Stage 3 — Iron deficiency anaemia: Haemoglobin falls. Microcytic hypochromic red cells (MCV <80 fL, MCH <27 pg). Pencil cells, target cells on film. Raised RDW (anisocytosis).

Biochemical diagnosis panel:
- Haemoglobin + RBC indices (MCV, MCH, MCHC)
- Serum ferritin (↓ in IDA, ↑ in ACD/inflammation)
- Serum iron (↓)
- Total Iron Binding Capacity — TIBC (↑ in IDA, ↓ in ACD)
- Transferrin saturation = serum iron ÷ TIBC × 100 (↓ in IDA)
- Reticulocyte haemoglobin content (RHC) — early, sensitive marker

India burden: ~50% of pregnant women, 20% of non-pregnant women, and 7% of men are iron-deficient. Causes: inadequate dietary intake (low meat in vegetarian diets), high phytate diet reducing absorption, chronic parasitic infestations (hookworm — significant in rural India, causes GI blood loss).

Government programme: Anaemia Mukt Bharat — weekly iron-folic acid supplementation for adolescents, daily supplementation in pregnancy.

CLINICAL PEARL

Ferritin as an acute-phase reactant: Ferritin rises in inflammation, infection, liver disease, and malignancy — independent of iron stores. Therefore, a "normal" or "high" ferritin does NOT rule out iron deficiency in a patient with active infection or chronic disease. In such patients, transferrin saturation <20% and reticulocyte haemoglobin <28 pg are more reliable markers of functional iron deficiency. This distinction between true iron deficiency and anaemia of chronic disease (ACD) is clinically critical — ACD responds to treating the underlying condition, not iron therapy.

Calcium, Phosphorus, and Bone Mineral Metabolism

Calcium — the most abundant mineral in the body (~1 kg; 99% in bone as hydroxyapatite).

Calcium, Phosphorus, and Bone Mineral Metabolism

Figure: Calcium, Phosphorus, and Bone Mineral Metabolism

Multi-panel illustration of calcium metabolism: blood calcium distribution and albumin correction, PTH-calcitriol-calcitonin regulatory axis, phosphorus metabolism with Ca×PO4 product, and osteoporosis pathology with DEXA diagnosis

Blood calcium (total): 8.5–10.5 mg/dL
- ~50% ionised Ca²⁺ (physiologically active)
- ~40% protein-bound (mainly to albumin — low albumin → low total Ca but normal ionised Ca)
- ~10% complexed with citrate, phosphate

Calcium regulation (triple axis: PTH, Calcitriol, Calcitonin):
- ↓ serum Ca²⁺ → PTH released from parathyroid → ↑ renal Ca reabsorption, ↑ bone resorption, ↑ renal 1α-hydroxylase → ↑ calcitriol → ↑ intestinal Ca absorption → Ca²⁺ rises
- Calcitonin (from C-cells of thyroid) → opposes PTH → inhibits osteoclasts → ↓ Ca

Hypocalcaemia: paraesthesiae, muscle cramps, tetany (carpal spasm), positive Chvostek sign (tapping facial nerve → facial twitch), Trousseau sign (BP cuff inflation → carpal spasm). Causes: hypoparathyroidism, Vitamin D deficiency, renal failure.

Hypercalcaemia: "bones, stones, groans, psychic moans" — bone pain, renal stones (calcium oxalate), abdominal pain, confusion. Causes: primary hyperparathyroidism (commonest), malignancy (bone metastases, PTHrP), hypervitaminosis D.

Phosphate: 85% in bone, 15% intracellular. Reciprocal relationship with calcium (high phosphate → lowers calcium). Elevated in CKD (kidneys cannot excrete phosphate → secondary hyperparathyroidism → renal osteodystrophy).